Zyra's plants can attack Akali in her new shroud when the plants are given true sight
As shown in the video Akali is not "obscured" to Zyra's plants when the plants are given true sight of Akali. The turrets give the plants true sight of Akali which then allows the plants to attack her but the turret themselves cannot attack Akali.
This also applies when plants are given true sight of Akali through the oracle's extract on the howling abyss (where I actually found this bug)
Skins and items do not affect the outcome and was reproduced 100% of the time, all you needed was a Zyra's plants with some form of true sight near the akali.
Other AI based targeting systems in the game (Heimerdinger turrets) cannot attack Akali while she obscured and within true sight.